ἤϊσαν

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ὅσα ἦν νενοσσευμένα ὀρνίθων γένεα → as many species of birds as had their nests, all the other kinds of birds which had been hatched

Source

French (Bailly abrégé)

3ᵉ pl. impf. épq. et ion. de εἶμι.

Russian (Dvoretsky)

ἤϊσαν: эп.-ион. (= ᾖεσαν) 3 л. pl. impf. к εἶμι.